Click on the Target 1 bone. This should create a channel in the pose. Right-clicking on that channel should give you the option to create a New Constraint. Create a Path constraint.In the property dialog, click the Object drop-down and select "Right Hand Control". Very likely (since there is only one spline in the Right Hand Control model) the program will automatically pick the right spline. If it doesnt, click the Eyedropper button and then click in the Pose window on the spline you created in the Hand Control object. Your bone should now adhere to the spline, at one end or another. Click the 'Translate Only' checkbox so that your bone doesn't orient along the spline (which can be distracting).
Why one endpoint over another? One of those endpoints is considered the start of the spline (at Ease zero) and the other one is considered the end of the spline (at Ease 100). I'm not really sure how the program chooses... probably the first CP you laid out when you were adding points. In any event, if it stuck to the wrong end, you can correct that by setting the Ease to the other end.
Now click on the Target 2 bone. Using the same procedure as above, assign it to the Path constraint. Change its Ease to "33" (or "67" if you had to set the Ease of the Target 1 bone to 100, above). This should move it to one-third of the way along the path. Similarly, assign the Target 3 bone to the 67 percent Ease point, and the Target 4 bone to the 100 percent Ease point. Now the bones should be distributed evenly across the Path.
Now, what was the point of all that? All will be revealed on the
next page.
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